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Does Dissolved Organic Matter Composition Help Explain the Concentrations of Bioavailable Macronutrients in Organic Matter-Rich Freshwaters?

Berggren, Martin LU ; Rulli, Mayra P.D. LU ; Bergström, Ann Kristin ; Sponseller, Ryan A. and Hensgens, Geert (2025) In Freshwater Biology 70(11).
Abstract

Dissolved organic matter (DOM) is a major source of macronutrients to freshwaters, yet it has variable and poorly understood bioavailability. Because intrinsic variation in bioavailability is caused by chemical structures of organic nutrients, DOM composition data should improve predictions of bioavailable resource pool sizes. We hypothesized that bioavailable organic carbon (C) and nitrogen (N) fractions are made up of freshly produced humic- and protein-like DOM, respectively, whereas bioavailable phosphorus (P) is linked to microbially-derived DOM with potential organophosphate content and/or to humic-like structures associated with DOM-Fe-phosphate complexes. These hypotheses were tested from surface waters collected at eight,... (More)

Dissolved organic matter (DOM) is a major source of macronutrients to freshwaters, yet it has variable and poorly understood bioavailability. Because intrinsic variation in bioavailability is caused by chemical structures of organic nutrients, DOM composition data should improve predictions of bioavailable resource pool sizes. We hypothesized that bioavailable organic carbon (C) and nitrogen (N) fractions are made up of freshly produced humic- and protein-like DOM, respectively, whereas bioavailable phosphorus (P) is linked to microbially-derived DOM with potential organophosphate content and/or to humic-like structures associated with DOM-Fe-phosphate complexes. These hypotheses were tested from surface waters collected at eight, unproductive and organic matter-rich stream and lake sites, from which we performed C, N and P microbial bioassays with flow cytometry in combination with analyses of DOM composition using fluorescence excitation-emission matrix (EEM) analysis. Bioavailable C followed the predicted patterns, with strong links to fluorescent features indicating recently produced DOM. Surprisingly, bioavailable N was relatively poorly related to DOM composition, including protein-like fluorescence, and was instead driven mainly by the amount of inorganic N. The bioavailable P showed patterns in support of the hypothesized link to microbially-derived organic components, whereas its relationships to free or complex-bound forms of inorganic phosphate were inconclusive. Thus, the strength of the hypothesized patterns varied. Nonetheless, in addition to the variability in bioavailable nutrient concentrations explained by standard bulk nutrient variables, we show that DOM composition variables made significant and unique contributions to explaining the variance in bioavailable C (19%), N (13%) and P (18%). Therefore, improved regression models for bioavailable nutrient concentrations could be achieved by including DOM composition among the explanatory variables. Overall, DOM composition analysis is a promising tool to improve prediction and develop our understanding of bioavailable macronutrients in organic matter-rich freshwaters.

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author
; ; ; and
organization
publishing date
type
Contribution to journal
publication status
published
subject
keywords
bioavailability, dissolved organic matter, fluorescence excitation-emission matrix analysis, macronutrients
in
Freshwater Biology
volume
70
issue
11
article number
e70141
publisher
Wiley-Blackwell
external identifiers
  • scopus:105022700578
ISSN
0046-5070
DOI
10.1111/fwb.70141
language
English
LU publication?
yes
additional info
Publisher Copyright: © 2025 The Author(s). Freshwater Biology published by John Wiley & Sons Ltd.
id
1270eeff-b861-4971-a69b-e5d4767d466f
date added to LUP
2026-01-22 14:18:48
date last changed
2026-01-23 09:12:42
@article{1270eeff-b861-4971-a69b-e5d4767d466f,
  abstract     = {{<p>Dissolved organic matter (DOM) is a major source of macronutrients to freshwaters, yet it has variable and poorly understood bioavailability. Because intrinsic variation in bioavailability is caused by chemical structures of organic nutrients, DOM composition data should improve predictions of bioavailable resource pool sizes. We hypothesized that bioavailable organic carbon (C) and nitrogen (N) fractions are made up of freshly produced humic- and protein-like DOM, respectively, whereas bioavailable phosphorus (P) is linked to microbially-derived DOM with potential organophosphate content and/or to humic-like structures associated with DOM-Fe-phosphate complexes. These hypotheses were tested from surface waters collected at eight, unproductive and organic matter-rich stream and lake sites, from which we performed C, N and P microbial bioassays with flow cytometry in combination with analyses of DOM composition using fluorescence excitation-emission matrix (EEM) analysis. Bioavailable C followed the predicted patterns, with strong links to fluorescent features indicating recently produced DOM. Surprisingly, bioavailable N was relatively poorly related to DOM composition, including protein-like fluorescence, and was instead driven mainly by the amount of inorganic N. The bioavailable P showed patterns in support of the hypothesized link to microbially-derived organic components, whereas its relationships to free or complex-bound forms of inorganic phosphate were inconclusive. Thus, the strength of the hypothesized patterns varied. Nonetheless, in addition to the variability in bioavailable nutrient concentrations explained by standard bulk nutrient variables, we show that DOM composition variables made significant and unique contributions to explaining the variance in bioavailable C (19%), N (13%) and P (18%). Therefore, improved regression models for bioavailable nutrient concentrations could be achieved by including DOM composition among the explanatory variables. Overall, DOM composition analysis is a promising tool to improve prediction and develop our understanding of bioavailable macronutrients in organic matter-rich freshwaters.</p>}},
  author       = {{Berggren, Martin and Rulli, Mayra P.D. and Bergström, Ann Kristin and Sponseller, Ryan A. and Hensgens, Geert}},
  issn         = {{0046-5070}},
  keywords     = {{bioavailability; dissolved organic matter; fluorescence excitation-emission matrix analysis; macronutrients}},
  language     = {{eng}},
  number       = {{11}},
  publisher    = {{Wiley-Blackwell}},
  series       = {{Freshwater Biology}},
  title        = {{Does Dissolved Organic Matter Composition Help Explain the Concentrations of Bioavailable Macronutrients in Organic Matter-Rich Freshwaters?}},
  url          = {{http://dx.doi.org/10.1111/fwb.70141}},
  doi          = {{10.1111/fwb.70141}},
  volume       = {{70}},
  year         = {{2025}},
}