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Digital Microfluidics-Driven Cell-Free Protein Synthesis Platform Reveals Expression and Stability Determinants for Phytoglobins and Cysteine-to-Alanine Substituted Variants

Groth, Leonard LU and Bülow, Leif LU (2025) In Antioxidants 14(11).
Abstract

Heme proteins are central to metabolism and stress responses but remain challenging to express recombinantly due to cytotoxicity and folding constraints. Phytoglobins (Pgbs) exemplify these difficulties, as expression protocols often fail to translate across protein species. Here, we used a cell-free protein synthesis (CFPS) platform powered by digital microfluidics to screen expression determinants for sugar beet Pgb 1.2 (BvPgb 1.2), its C86A variant, and three of eight newly identified oat Pgbs (AsPgbs), including their cysteine-to-alanine substituted variants. Benchmarking with multiple solubility tags and cell-free blends revealed protein- and variant-specific preferences, with alanine substitutions frequently improving expression... (More)

Heme proteins are central to metabolism and stress responses but remain challenging to express recombinantly due to cytotoxicity and folding constraints. Phytoglobins (Pgbs) exemplify these difficulties, as expression protocols often fail to translate across protein species. Here, we used a cell-free protein synthesis (CFPS) platform powered by digital microfluidics to screen expression determinants for sugar beet Pgb 1.2 (BvPgb 1.2), its C86A variant, and three of eight newly identified oat Pgbs (AsPgbs), including their cysteine-to-alanine substituted variants. Benchmarking with multiple solubility tags and cell-free blends revealed protein- and variant-specific preferences, with alanine substitutions frequently improving expression and purification yields. Oxidative additives such as glutathione disulfide, alone or combined with protein disulfide isomerase, consistently enhanced production, underscoring the importance of redox environments for Pgb stability. Two selected variants were scaled up and yielded putative soluble apo-form proteins. The results highlight how CFPS enables rapid, parallelized identification of expression requirements while uncovering the role of conserved cysteines and redox conditions in Pgb biogenesis.

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author
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organization
publishing date
type
Contribution to journal
publication status
published
subject
keywords
cell-free protein synthesis, cysteine, digital microfluidics, expression determinants, heme protein, phytoglobin, redox environment, solubility tag screen
in
Antioxidants
volume
14
issue
11
article number
1317
publisher
MDPI AG
external identifiers
  • scopus:105023156145
  • pmid:41300473
ISSN
2076-3921
DOI
10.3390/antiox14111317
language
English
LU publication?
yes
additional info
Publisher Copyright: © 2025 by the authors.
id
b88fcf8e-975f-4aef-b8e7-daa1d18899a5
date added to LUP
2026-01-22 13:30:35
date last changed
2026-01-23 03:00:13
@article{b88fcf8e-975f-4aef-b8e7-daa1d18899a5,
  abstract     = {{<p>Heme proteins are central to metabolism and stress responses but remain challenging to express recombinantly due to cytotoxicity and folding constraints. Phytoglobins (Pgbs) exemplify these difficulties, as expression protocols often fail to translate across protein species. Here, we used a cell-free protein synthesis (CFPS) platform powered by digital microfluidics to screen expression determinants for sugar beet Pgb 1.2 (BvPgb 1.2), its C86A variant, and three of eight newly identified oat Pgbs (AsPgbs), including their cysteine-to-alanine substituted variants. Benchmarking with multiple solubility tags and cell-free blends revealed protein- and variant-specific preferences, with alanine substitutions frequently improving expression and purification yields. Oxidative additives such as glutathione disulfide, alone or combined with protein disulfide isomerase, consistently enhanced production, underscoring the importance of redox environments for Pgb stability. Two selected variants were scaled up and yielded putative soluble apo-form proteins. The results highlight how CFPS enables rapid, parallelized identification of expression requirements while uncovering the role of conserved cysteines and redox conditions in Pgb biogenesis.</p>}},
  author       = {{Groth, Leonard and Bülow, Leif}},
  issn         = {{2076-3921}},
  keywords     = {{cell-free protein synthesis; cysteine; digital microfluidics; expression determinants; heme protein; phytoglobin; redox environment; solubility tag screen}},
  language     = {{eng}},
  number       = {{11}},
  publisher    = {{MDPI AG}},
  series       = {{Antioxidants}},
  title        = {{Digital Microfluidics-Driven Cell-Free Protein Synthesis Platform Reveals Expression and Stability Determinants for Phytoglobins and Cysteine-to-Alanine Substituted Variants}},
  url          = {{http://dx.doi.org/10.3390/antiox14111317}},
  doi          = {{10.3390/antiox14111317}},
  volume       = {{14}},
  year         = {{2025}},
}