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A histidine pseudokinase modulates polar growth and cell shape in Streptomyces venezuelae

Singh Mavi, Parminder LU and Flärdh, Klas LU (2026) In Communications Biology 9(1).
Abstract

Polar growth and cell shape determination in mycelium-forming Streptomyces bacteria depends on the function of a polarly localised multiprotein complex that directs cell wall synthesis – the polarisome. This complex assembles around the essential cell polarity determinant DivIVA, alongside other largely unknown components. We report here the discovery of a conserved hybrid histidine kinase-like protein, PsmA, that interacts and co-localises with DivIVA at the hyphal tips. Deletion of psmA affects the shape and dynamics of the polarisome, leading to aberrant cell shape and hyphal hyperbranching. PsmA is a pseudokinase that lacks the critical histidine residue in its catalytic core. Our results suggest that PsmA tunes the dynamics and... (More)

Polar growth and cell shape determination in mycelium-forming Streptomyces bacteria depends on the function of a polarly localised multiprotein complex that directs cell wall synthesis – the polarisome. This complex assembles around the essential cell polarity determinant DivIVA, alongside other largely unknown components. We report here the discovery of a conserved hybrid histidine kinase-like protein, PsmA, that interacts and co-localises with DivIVA at the hyphal tips. Deletion of psmA affects the shape and dynamics of the polarisome, leading to aberrant cell shape and hyphal hyperbranching. PsmA is a pseudokinase that lacks the critical histidine residue in its catalytic core. Our results suggest that PsmA tunes the dynamics and properties of the DivIVA-based polar organelle in streptomycetes in parallel to but not redundantly with Scy and FilP, two coiled-coil proteins known to influence polarisome properties. In summary, PsmA interacts with DivIVA and modulates the integrity of the growth zones at hyphal tips.

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author
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type
Contribution to journal
publication status
published
subject
in
Communications Biology
volume
9
issue
1
article number
345
publisher
Nature Publishing Group
external identifiers
  • scopus:105032234460
  • pmid:41620559
ISSN
2399-3642
DOI
10.1038/s42003-026-09620-z
language
English
LU publication?
yes
id
b541e340-2f93-44dd-9060-66a459a4b37d
date added to LUP
2026-04-23 12:53:04
date last changed
2026-06-04 15:37:48
@article{b541e340-2f93-44dd-9060-66a459a4b37d,
  abstract     = {{<p>Polar growth and cell shape determination in mycelium-forming Streptomyces bacteria depends on the function of a polarly localised multiprotein complex that directs cell wall synthesis – the polarisome. This complex assembles around the essential cell polarity determinant DivIVA, alongside other largely unknown components. We report here the discovery of a conserved hybrid histidine kinase-like protein, PsmA, that interacts and co-localises with DivIVA at the hyphal tips. Deletion of psmA affects the shape and dynamics of the polarisome, leading to aberrant cell shape and hyphal hyperbranching. PsmA is a pseudokinase that lacks the critical histidine residue in its catalytic core. Our results suggest that PsmA tunes the dynamics and properties of the DivIVA-based polar organelle in streptomycetes in parallel to but not redundantly with Scy and FilP, two coiled-coil proteins known to influence polarisome properties. In summary, PsmA interacts with DivIVA and modulates the integrity of the growth zones at hyphal tips.</p>}},
  author       = {{Singh Mavi, Parminder and Flärdh, Klas}},
  issn         = {{2399-3642}},
  language     = {{eng}},
  number       = {{1}},
  publisher    = {{Nature Publishing Group}},
  series       = {{Communications Biology}},
  title        = {{A histidine pseudokinase modulates polar growth and cell shape in Streptomyces venezuelae}},
  url          = {{http://dx.doi.org/10.1038/s42003-026-09620-z}},
  doi          = {{10.1038/s42003-026-09620-z}},
  volume       = {{9}},
  year         = {{2026}},
}